Cork Cambium
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Cork cambium (pl. cambia or cambiums) is a tissue found in many vascular plants as a part of the epidermis. It is one of the many layers of bark, between the cork and primary
phloem Phloem (, ) is the living biological tissue, tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as ''photosynthates'', in particular the sugar sucrose, to the rest of the plant. This tran ...
. The cork cambium is a lateral
meristem The meristem is a type of tissue found in plants. It consists of undifferentiated cells (meristematic cells) capable of cell division. Cells in the meristem can develop into all the other tissues and organs that occur in plants. These cells conti ...
and is responsible for secondary growth that replaces the epidermis in
root In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the su ...
s and stems. It is found in woody and many herbaceous
dicot The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, t ...
s,
gymnosperm The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, ''Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, Î³Ï…Î¼Î½ÏŒÏ ...
s and some
monocot Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one Embryo#Plant embryos, embryonic leaf, or cotyledon. Th ...
s (monocots usually lack secondary growth). It is one of the plant's meristems – the series of tissues consisting of embryonic disk (incompletely differentiated) cells from which the plant grows. The function of cork cambium is to produce the
cork Cork or CORK may refer to: Materials * Cork (material), an impermeable buoyant plant product ** Cork (plug), a cylindrical or conical object used to seal a container ***Wine cork Places Ireland * Cork (city) ** Metropolitan Cork, also known as G ...
, a tough protective material.Junikka, L. (1994) "Macroscopic bark terminology". ''IAWA Journal'' 15(1): 3–45Trockenbrodt, M. (1990) "Survey and discussion of the terminology used in bark anatomy". ''IAWA Bulletin, New Series'' 11: 141–166 Synonyms for cork cambium are bark cambium, pericambium and phellogen. Phellogen is defined as the meristematic cell layer responsible for the development of the periderm. Cells that grow inwards from there are termed ''phelloderm'', and cells that develop outwards are termed ''phellem'' or cork (note similarity with
vascular cambium The vascular cambium is the main growth tissue in the stems and roots of many plants, specifically in dicots such as buttercups and oak trees, gymnosperms such as pine trees, as well as in certain other vascular plants. It produces secondary xyle ...
). The
periderm Bark is the outermost layers of stems and roots of woody plants. Plants with bark include trees, woody vines, and shrubs. Bark refers to all the tissues outside the vascular cambium and is a nontechnical term. It overlays the wood and consist ...
thus consists of three different layers: * phelloderm – inside of cork cambium; composed of living parenchyma cells * phellogen (cork cambium) – meristem that gives rise to periderm * phellem (cork) – dead at maturity; air-filled ery variable between different species, and is also highly dependent on age and growth conditions, as can be observed from the different surfaces of bark, which may be smooth, fissured, tesselated, scaly, or flaking off.


Economic importance

Commercial
cork Cork or CORK may refer to: Materials * Cork (material), an impermeable buoyant plant product ** Cork (plug), a cylindrical or conical object used to seal a container ***Wine cork Places Ireland * Cork (city) ** Metropolitan Cork, also known as G ...
is derived from the bark of the
cork oak ''Quercus suber'', commonly called the cork oak, is a medium-sized, evergreen oak tree in the section ''Quercus'' sect. ''Cerris''. It is the primary source of cork for wine bottle stoppers and other uses, such as cork flooring and as the core ...
''(Quercus suber).'' Cork has many uses including wine bottle stoppers, bulletin boards, coasters, hot pads to protect tables from hot pans, insulation, sealing for lids, flooring, gaskets for engines, fishing bobbers, handles for fishing rods and tennis rackets, etc. It is also a high strength-to-weight/cost ablative material for aerodynamic prototypes in wind tunnels, as well as satellite launch vehicle payload fairings, reentry surfaces, and compression joints in thrust-vectored solid rocket motor nozzles. Many types of bark are used as
mulch A mulch is a layer of material applied to the surface of soil. Reasons for applying mulch include conservation of soil moisture, improving fertility and health of the soil, reducing weed growth and enhancing the visual appeal of the area. A mu ...
.


See also

*
Complementary cells Complementary cells are a mass of cells in plants, formed from the cork cambium at the position of the lenticels. It is a group of loosely arranged cells that aid in gaseous exchange through cork Cork or CORK may refer to: Materials * Cork (mat ...
*
Frost crack Frost crack or Southwest canker is a form of tree bark damage sometimes found on thin barked trees, visible as vertical fractures on the southerly facing surfaces of tree trunks. Frost crack is distinct from sun scald and sun crack and physically ...
* Sun scald (flora)


References

{{Biological tissue Plant anatomy